这里用的是倒装
主语：the sex ratios
定语： in certain parasitic wasp species that have a large excess of females.
谓语：are Immediately relevant to game theory
没有倒装的语序为：
the sex ratios in certain parasitic wasp species that have a large excess of females are Immediately relevant to game theory .
在某些雄性黄蜂占多数的黄蜂种类中，其性别比列直接涉及到了博弈论。
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